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Lichen

Messages from the 1970s…

I’ve recently been browsing a set of reports that Francis Rose wrote for the Nature Conservancy Council in the early 1970s. Mainly, but not exclusively, about lichens, they contain a wealth of information about the visits he (with others including Brian Coppins and our own Russell Gomm) made to Cumbria’s woodlands in search of epiphytes. They are fascinating reads, in oh-so-many ways.

Firstly, there’s the human stuff: there was a lot of searching and lichen bothering going on, in fair weather and foul. The main (1971) report covers 24 days during which he visited 75 sites plus additional trees, not to mention the areas that were looked at but didn’t make the cut. Many of the woods that he identified as being the best in Lakeland are still regarded as such: Seathwaite, Yew Crag, Scales Wood, Naddle. Some that are now recognised as being very important, like Rydal, didn’t get the attention they maybe deserved.

The reports give a marvellous sense of an understanding being developed. The factors influencing which woodlands are good for which lichens in Cumbria are being observed and weighed: the indices of ecological continuity are on their way to being sorted. Sites in drier, sheltered areas (with basic barked trees) tend to have a different (and possibly more species-rich) lichen flora than more upland, wetter woodlands where the nutrients have presumably been leached from the bark. These days, the differences are recognised in the Southern Oceanic Woodland Index and the Upland Rainforest Index.

Lobaria pulmonaria near Rydal
Lobaria pulmonaria near Rydal

But woodland history and management override other factors: young/ coppiced woodlands have more limited epiphyte populations. And then there’s air pollution: the effects of coal burning and sulphur dioxide production were much more obvious then. The northern and eastern Lakes, more sheltered from the effects of industrial pollution, had retained better lichen floras. There’s also the soil: not only do more basic soils allow ash and elm to grow, but oaks on more basic soils tend to have richer lichen floras. And as soon as one starts to climb through a wood, the effects of leaching are visible: the higher parts of wood have a more “upland, acid bark loving, flora”.

All of which hold true today, but when it comes to the species, the reports can read as though they are talking about somewhere completely different. Obviously, many of the species names have changed; many more lichens are now recognised and described. But Parmotrema perlatum was only found at one site in the Windermere/ Coniston/Ambleside area (these days it is prolific) and Normandina pulchella is noteworthy as an indicator of relict Lobarion: how some things have bounced back! Maybe, to look on the bright side, there are more lichens now in many places.

Parmotrema perlatum near Ambleside
Parmotrema perlatum near Ambleside

Another example of how things have changed is the description of Physcia aipolia as “interesting”. It is, these days, prolific on my soft fruit bushes. The Xanthorion is held to be an interesting community: the prolific growths of nitrophilous lichens that we see today are far from being an issue. Lowther Park is described as being “probably the richest so far discovered” in Northern England. My explorations there suggest a lot of nitrogen enrichment, and the loss of all the interesting lichens. One report gives details of individual trees: it will be worth revisiting these to see what exactly is there today.

Physcia aipolia near Ambleside
Physcia aipolia near Ambleside
Evernia prunastri at  Lowther Park
Evernia prunastri at Lowther Park

Overall, it is probably the losses that stand out more than the gains: Great Wood, Borrowdale, was described as having the three Lobaria species in an “abundance unparalleled in Northern England”. Whilst some (of all three) remains, the elms are gone and the decline in Lobarion well known. Across Lakeland, these excursions came across large old trees that retained lichens from pre-industrial times; wayside oaks with Sticta limbata, Lobaria virens and Nephroma. Fifty years on and they aren’t on the roadsides, though some of the old ash pollards have them just about hanging on.

Lobaria pulmonaria, L. amplissima and L. virens in Great Wood
Lobaria pulmonaria, L. amplissima and L. virens in Great Wood

On the other hand, some things remain the same: the “abundance” of Bryoria fuscescens in Holme Wood, Loweswater, that struck Francis Rose is my main memory from our 2020 visit. And then there’s the things that were found that need searching for again: Usnea ceratina in Pull Garth Woods; Heterodermia obscurata at Lodore. If these reports do one thing, they urge us to get outside with a hand lens. Cue the suggestions for next winter’s programme!

Bryoria fuscescens at Holme Wood
Bryoria fuscescens at Holme Wood

And finally, it is interesting to note the report’s recommendations: that many of the sites in Borrowdale should be “given National Nature Reserve status within an envelope”, and other sites given protection too. Fifty-plus years on, it may be that the former is being considered. Was that the sound of a stable door I just heard?

Text and photos: Pete Martin

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Lichen Trip report

Brown Cove Lichen Report 11 June 2023

Participants: Geoffrey Haigh, Raymond and Sue Griffiths, Pete Martin, Caz Walker, Chris Cant, Jack Tomlinson, Clare Shaw and reporter Peter Bisset

After a long spell of hot dry weather, things looked more promising for the eight lichenologists than the solitary bryologist. Clare Shaw searching along the beck and tarn has already reported on her findings and pointed out some to us. The blazing sun and steady climb dictated even more slowly moving exploration than normal. What a contrast to the Naddle meet, how lucky we are to live in a country with such distinct seasons. I followed Clare up the beck admiring the colourful mounds of bryophytes with little clue as to what I was looking at and came across some Peltigera, I called it out to Caz who accepted my membranacea determination without climbing down to me. Looking at the photographs later it was clear that this was mixed with P. praetextata, a new record for the monad.

Keppel Cove dam
Keppel Cove dam

Brown Cove has an interesting base rich chemistry, but being shallow with plenty of green algae – not appealing for a wild swim. Known as the only UK site for Dermatocarpon deminuens and intensively surveyed by Alan Orange the monad is well recorded. However, it was good checking what could still be found. The intermittently inundated shore line with the water level now very low made it easier to spot the Dermatocarpon, jelly lichens and Ionapsis lacustris. D. luridum and D. intestiniforme could be distinguished by the former going green on wetting, but any others need lab work to be sure. Pete pointed out Lathagrium (Collema) dichotomum on the rocks, a species normally found under water.

An extended lunch break is always good for looking at the lichens on the rocks under your legs, a dropped sandwich often extends the species list, Lecanora soralifera was one of these.

Being a somewhat superficial lichenologist I like the showy or unusual, Umbilicaria cylindrica and Ophioparma ventosa provided the former. Agonimia tristicula one of the latter. However, the high point of my day was when the track crossed the corner of a new monad on the way back down. Peering into a sandy/peaty hole to contribute to the quick assessment, an unusual looking light green lobate species in amongst the Micarea lignaria had me opening page 44 of Dobson (7th edition), generic keys – ‘foliose’ but I wasn’t getting anywhere. Caz immediately identified it as Baeomyces placophyllus, new to me and saving me a frustrating time in the keys where the genus is ‘crustose’.

Baeomyces placophyllus
Baeomyces placophyllus

It was a hot day for a hike so thanks for arranging parking at Greenside. A fitness event in Glenridding would in any case have made parking impossible at any price!

Text and photos: Peter Bisset

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Lichen Trip report

Burnbanks trip report 18-5-2023

Ten of us gathered on a sunny May morning to look at lichens. We crossed the road, went through a gate and there was a slope. Small outcrops beckoned. The bracken wasn’t yet tall enough to be in the way. Yellowhammers wheezed about no cheese. This was Caz and Chris’s local patch, and they had an itinerary planned…

Outcrop one had some very impressive Massalongia carnosa: it was strange to see it dry. There were lichens to explain to first timers: Rhizocarpon geographicum; a conspiracy of Cladonias; Ochrolechia androgyna to demonstrate a C+ red reaction on; Pertusaria corallina for K+ yellow; Ephebe lanata to stroke.

Heading across the slope (and edging a little bit higher) we came to an outcrop where Chris and Caz had previously found Lecidea fuliginosa. It’s not in Dobson, so there had been some uncertainty. I’d not seen it before: it’s an interesting brown nobbly crust, with darker black-brown apothecia. My photos suggest the lichen equivalent of home-made chocolate chip cookies, but maybe I’m taking the food analogies too far. There was also a Small Copper butterfly, interesting little still-lifes of Cladonia furcata and very fertile Xanthoparmelia conspersa; a good example of Lecanora sulphurea taking over (i.e. parasitising the algae of) another lichen. Was it Tephromela atra or Lecanora gangaleoides? I must admit I didn’t check.

There was Ochroclechia tartarea and parella to compare; Diploschistes scruposus hiding yellow tinged behind an outcrop, and Lecidea grisella’s cracks to ponder. We found some shiny Cetraria aculeata too.

Lunch was taken in the shelter of a little quarry, which occupied us for a while afterwards. There was Opegrapha zonata and O.gyrocarpa (now renamed Enterographa zonata and Gyrographa gyrocarpa respectively) to discuss, and some evidence of metals in the rock: Tremolecia atrata looked rusty and there were Stereocaulons aplenty. I’d not tried the KC test on S. evolutum before: it gives a fleeting “violet”. Well, a different colour from the C and the K anyway.

A little further uphill and there was a dramatic boulder: Bryoria fuscescens cascaded for several feet down the lower side. There was Tuckermanopsis chlorophylla to compare with nearby Platismatia glauca, hi-vis Arthrorhaphis citrinella and an intriguing Lecanora with pearly pruinose apothecia: C-, K+ yellow, P+ orange. There was a suggestion that this was Lecanora subcarnea, but the chemistry didn’t agree with Dobson. Though it looked like the picture. The puzzlement was resolved back at base when it transpired that the chemistry listed in LGBI3 agreed with our “in the field” records: result! Another new one for me.

Over the top of the next knoll was a little area of heath. Hands and knees alternated with standing and staring at our toes. Apart from Cladonia we found a couple of pieces of Cetraria islandica: cue a debate about pronunciation. And a small population of Cetraria muricata: much more spiny, closer together, pseudocyphellae not in pits.

A short wander took us to the slope overlooking Haweswater and the green expanse of Naddle woods. The Willow Warbler chorus that had been with us all day continued. There was Protoparmelia badia on the final erratic, a last chance of the day to say “ornithocoprophilous” (whilst looking at probable Xanthoria ucrainica) and a great expanse of Parmelia omphalodes, split by a line of bilberry.

Thank you everyone for coming, Caz and Chris for the planning, and the weather gods for being kind. All in all it was a very good start to the rocky summer season (as opposed to the winter woods).  

Text: Pete Martin
Photos: Chris Cant, John Adams, Pete Martin

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Lichen Trip report

Seathwaite trip report 22-4-2023

We were going to the rainforest so… it rained. Which put some folk off. But it didn’t rain that much: the four of us who gathered to look at lichen ended up being able to look at things without too much watery difficulty.

Predictably, we got rather distracted before the main event. The moment we left the road at Seathwaite Bridge we found a wall jam-packed with typical saxicolous species: Parmelia saxatilis; Xanthoparmelia conspersa; Lecanora soralifera; Hypotrachyna revoluta sensu lato; Ochrolechia androgyna. So it was a while before we were heading up the slope from tree to tree, noting the difference between the lichens on older oaks and those on hazels, though some species like Thelotrema lepadinum seemed happy on both.

It was, of course, the Lobarion lichens on the older ashes that stole the show. We were examining a Collema (subflaccidum I think) when we realised there was a profuse growth of Peltigera praetextata on the other side of the tree, which led us to notice the sorediate margins of Nephroma parile. The next tree had profuse Nephroma laevigatum for contrast, with apothecia the other way round from those on Peltigera. There was Sticta fuliginosa too, which had us sniffing our fingers for fish. At some point someone said “the more knackered the tree, the better the lichens” and I couldn’t disagree.

There’s several curiosities about these Seathwaite woods. The ashes aren’t that big, but are very decrepit and have amazing lichens in spite of it being a mine and industrial site just a couple of hundred years ago. The Borrowdale yews, where we had lunch, are pretty enigmatic too. And there’s no Lobaria pulmonaria there. Except there is now, because of the translocations undertaken in 2020. We looked at a few of these: some were growing really healthily.

The translocations of dual morph Sticta canariensis were doing OK too: the brown lobes have cyanobacteria (“blue-green algae”) as the photobiont, whilst the green lobes that protrude from them have algae doing the job. The fungi is the same in both parts. We quickly found a larger, “free-living” population of dual morph S. canariensis, and then some large thalli of vivid green Ricasolia (was Lobaria) virens, complete with little volcanoes on the lobes and beautiful apothecia.

We made out way back through the wood, hopping from oak to ash to hazel but finally finished at an ash with the largest growth of Sticta I’ve ever seen. There was more than I could shake my stick at; great overlapping brackets cascaded down the trunk. I think it is Sticta sylvatica, but I struggle with telling the difference between that and S. fuliginosa.

I say finished, but of course we didn’t. There was a little revision on the wall at the entrance to the wood, and then a quick look at the Candelaria concolor on the rowans in the car park. Very different in scale, habitat requirements and conservation evaluation. There seemed rather more than I remembered. A nice little thing, even if it’s not as dramatic as the Lobarion.

Text and photos: Pete Martin

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Lichen Trip report

Hutton Roof lichen trip report 15-3-23

Ten folk braved the March snow to look at lichens near Hutton Roof this week. And there were half a dozen for the bryo option: this looking at small things is getting popular! Right on the wall by the meeting point in the village was Diploschistes muscorum, a crusty species that has the white-with-a-yellow-tinge crust of its commoner cousin Diploschistes scruposus, and the hollowed out apothecia too. But D. muscorum is parasitic on Cladonias (particularly on limestone in my experience). It seems to smother them, covering the podetia in a white blanket. And then eat them up before moving on to the mosses.

A short distance up the lane we found a limestone wall to occupy us for half an hour or so: Caloplaca flavescens,  Diploicia canescens, Acrocordia conoidea and other typical species. The odd silicious rock had Parmelia saxatilis, Rhizocarpon geographicum and Opegrapha gyrocarpa to make them stand out. And then it was on to the Ash trees: with a lot of Physcia adscendens and Lecidella elaeochroma to discuss. Adjacent Punctelia subrudecta and Parmelia sulcata thalli gave good illustrations of their different pseudocyphellae: points versus lines. Bubbling Curlews and screeching Jays provided an interesting soundtrack.

And then there was an outcrop: Dermatocarpon miniatum as big as (small) elephants’ ears; bubbling thalli of Collema (now Lathagrium) auriforme and an unfamiliar tiny jelly lichen whose proportionally long extended lobes were pruinose at the tips. We discovered later that it is Scytinium (was Collema) fragile, confirmed by Brian Coppins. A well camouflaged moth was disturbed- initial investigations suggest Ectropis crepescularia – The Engrailed. Nearby walls had frilly Peltigera praetextata; the ash tree above had Pertusaria leioplaca and a couple of specimens – intriguing white apothecia, and one that turned out to be Gyalecta truncigena.

We pushed on through the Hazel woods. A real-life Coenognium (was Dimerella) lutea was compared with the front of Dobson. Pseudoschismatomma (was Opegrapha) rufescens was seen on an ash. We lunched on the limestone pavement of the Rakes, with Blue Moor Grass (Sesleria caerulea) at our feet and a snowy Ingleborough in view. Typical limestone outcrop species were quickly found:  Squamarina cartilaginea; Petractis clausa; Collema cristatum etc. Protoblastenias were discussed. Was that a colour change with K or not? Could it be P. lilacina?

Higher up on the outcrops we found some beautiful rosettes of Caloplaca aurantia, the flat lobe ends contrasting nicely with the nearby convex ones of Caloplaca flavescens. We debated saxicolous Opegraphas, found Dermatocarpon luridum in a karren (non-geomorphologists might call it a runnel) and demonstrated the K/UV purple reaction of Porpidia tuberculosa on an erratic. We failed to find Solorina saccata. Maybe next time.

It began to rain, so headed down, though were delayed for a while by a wall with Lecidella scabra, Baeomyces rufus, Leptogium gelatinosum and a mysterious greeny C+ red crust with small white features. Is it young Trapelia coarctata? Somewhat eerily, we emerged onto the road just as the bryo party arrived from the other direction: perfect timing for a post-trip debrief. We found 90 or so species, all learnt at least something, and there’s samples to occupy us over the next few days. But we have barely scratched the surface of Hutton Roof…

Text: Pete Martin. Photos: Caz Walker, Chris Cant, Geoffrey Haigh, Pete Martin

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Lichen

The lichens of Hadrian’s Wood and Tarraby Lane, 3rd Feb 2023

Caz and I recently ran an introduction to lichens session for the first year students on the Wildlife and Media B.A. at the University of Cumbria’s Brampton Road Campus, Carlisle. We walked initially along Tarraby Lane, lined with beech trees which cast a deep shade for much of the year. This makes it hard for lichens to grow, so leafy lichens were few here and very small, with crustose and leprose (powdery) species dominating. There was also evidence of nitrogen pollution based on which species were present or absent. The damp wood at the end of the lane had more light and was perhaps better sheltered from pollution, so there was a greater range of lichen growth forms and species seen, including the shocking pink lichenicolous fungus Illosporiopsis christiansenii, one of the fungi which specialise in living on/in particular lichens. This one’s preferred habitat is the lichen genus Physcia.

This is what the students said about the day, and their photos are below:

I loved learning more about lichen. I’ve been interested in fungi for years now but never really thought to look at their wonderful symbiotic existence as lichens. An alga, a bacterium and a fungus – it really is fascinating! Staring at the little blue-grey growths on the sides of trees was quite eye-opening. I’d never spent the time to notice how detailed they are. Some were bumpy with protruding cup-like structures, others flat so flat they looked like dust, on branches some stuck out like coral. What I found the most exciting was learning that there are minute coral-pink fungi growing on the surface of some of the lichen. Tiny little fungi piggybacking on another fellow fungus in a completely altered symbiotic state! Honestly, what’s not to love?

Nathan  Greening

I really enjoyed learning about lichen id, before I never really gave them much thought, but being able to identify key features of a habitat due to the massive variety of the group is a great fieldcraft skill, and can tell me things about the biodiversity which may not have been so obvious otherwise.

Daniel Peters

We really enjoyed it too: it got us to go and look at somewhere we otherwise wouldn’t (and make 80-odd records for the BLS database). I won’t forget the lichen-rich litter bin in front of the campus – see below! Thank you to Alex Playford, the course tutor, for organising it.

Pete Martin and Caz Walker

Photos by Daniel Peters

Photos by Issy Drake

Photos by Nathan Greening

Photo by Pete Martin

The lichen-rich litter bin in front of the campus
The lichen-rich litter bin in front of the campus

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Lichen Trip report

Borrowdale trip report 15-2-23

Lichens at Ashness Wood and Moss Mire, by Watendlath Beck

There was a good turn out – 10 people looking at lichens – despite the weather forecast for a wet morning. After a few heavy showers we had a dry afternoon which allowed for some good licheneering.

This is a National Trust site, part of the Lodore-Troutdale Woods SSSI which is designated for its upland acidic oak-birch woodland and forms part of the internationally important Borrowdale Woods complex.

Most of the participants were new to the group and, in some cases, new to lichens so we took a while to get out of the Surprise View car park where the trees had lovely displays of acidic bark species, showing a range of growth forms and other features. Once everyone had had a go with x10 hand lenses and got their eye in for the tiny structures we need to look at in order to start the ID process, we set off, pausing to examine oak and birch trees along the way.

At Moss Mire, beside Watendlath Beck, we stopped for lunch and spent a few hours looking closely at trees (corticolous lichens), riverine rocks, shaded upland rock outcrops (saxicolous species) and terricolous habitats (lichens growing on the ground). This was in the southern part of monad NY2618; we ventured briefly into NY2617 as there seemed to be no records for that square and wrote a short list. All our records will be submitted to the British Lichen Society.

Mostly the species we saw were from the acidic bark lichen communities including Bryoria fuscescens, Sphaerophorus globosus, Ochrolechia tartarea, Ochrolechia androgyna, Dimerella pineti and Micarea stipitata. The Ochrolechia species allowed a demonstration of a useful chemical colour change where a drop of C (bleach) goes red. Mycoblastus sanguinarius was also common with red pigment visible beneath black apothecia. However some of this may turn out to be Mycoblastus sanguinarioides, only recently found in Britain, and distinguished by internal crystals which show up in polarised light under a compound microscope. It needs to be confirmed, but well done to Pete for taking a specimen and investigating.

We also saw Thelotrema lepadinum, or barnacle lichen, which indicates good quality woodland, and black fuzz on mossy shaded rock which turned out to be two species, Cystocoleus ebeneus and Racodium rupestre. In a few spots twigs and small branches had blown down, allowing us to examine canopy lichens. There was a fair amount of head-scratching over assorted Cladonia species, in corticolous, saxicolous and terricolous habitats, with Pete finding cool Cladonia caespiticia showing fruiting bodies on short stalks. Other terricolous Cladonia were the richly-branching C. portentosa, C. arbuscula and C. ciliata and there were a couple of sightings of the interesting Trapeliopsis pseudogranulosa overgrowing decaying moss and vegetation with orange and yellow-green granules.

On the way back to the car park there were a few hazel trees which yielded some alkaline bark species, not seen before. Hazels were notable for their absence in the Moss Mire area, perhaps because the rock, therefore soil, is acidic. However there was very little tree regeneration there of any species, or indeed herbaceous flora, which is a sure sign of heavy grazing and, long term, leads to the death of the woodland. This is very much in the news now – how best to regenerate oceanic woodland in the UK? Deer fencing can mean no browsing at all which some lichenologists dislike as tall ground flora and eventually saplings adversely affect lichens on trunks by shading them out. Alternatively a site could be managed for light grazing which allows some natural tree regeneration – but this means major deer culling and only a small number of livestock which could be an effort to monitor etc.

This was a great day with lots of lichen chat, skill sharing, looking and learning.

Caz Walker. Photos: Chris Cant, Pete Martin, Ann Lingard

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Bryophyte Lichen Trip report

Naddle trip report 18-1-23

Once again, the weather forecast was for sub-zero temperatures, ice and snow.  After reassurance from Lee Schofield that the roads to Haweswater were “passable with care” it was decided that the trip could go ahead.  We were warmly welcomed by Lee and the rest of the team at the RSPB reserve, and several of them joined us for the day.  Since Kerry was the only bryologist to brave the weather, she joined the lichen team for a combined visit, which was very educational on both sides.

We started with a visit to the well-known Lobaria pulmonaria Ash tree across the stream from the farm.  Sadly it had lost a large limb a couple of years ago, but this had provided the material for many attempted translocations by Chris and Caz.  We visited several of these during the course of the day, and while it is still early days many of them seemed to be doing well.

The breeze in this area was very chilly, so we moved into the more sheltered Mirkside wood, of multi-stemmed hazel with Oak, Ash and Alder.  Here we saw Lee’s astonishing discovery of Hazel Glove Fungus, sadly now much eaten by slugs, and accompanied by large quantities of the Glue Fungus which it parasitises.  Mycology aside, there were very good assemblages of the typical smooth bark lichens such as Graphis, Arthonia etc., the other Lobaria tree was located and the RSPB team were introduced to the concept of lichen speed (measured in hours per tree). The epiphytic liverworts Frullania dilatata, Metzgeria furcata and Radula complanata were also abundant here. A large patch of Riccardia palmata was found on one of the few logs not covered in snow. 

At lunchtime sitting in a wet snowdrift seemed attractive enough, but after some discussion we decided to accept the RSPB’s offer of a warm room with a log fire, tea and coffee, chocolate biscuits and mince pies.   Annabel described the future plans for the reserve and for landscape restoration in the surrounding areas.  There is fantastic potential but it very much depends on the government sorting out the farm subsidy payments.

After lunch we moved to the sunlit side of the valley, where there is an area of wood-pasture with veteran Ash trees.  There were many good representatives of more basic-bark species, a highlight being Normandina acroglypta. The melting snow enabled Kerry to find more bryophytes too, including frequent cushions of Bartramia pomiformis and one patch of Porella arboris-vitae on the crags above the farm. 

Overall it was a most enjoyable day, and a site well worth another visit in better weather.  Very many thanks to the RSPB for their hospitality and enthusiasm.

John Adams and Kerry Milligan. Photos by John Adams, Chris Cant and Kerry Milligan

Octospora bryophilous fungus note below

Octospora bryophilous fungus

Caz found a distinctive fungus growing on Bryum moss on an ash tree. Chris found spores in a fruit, with 4 or more spores per ascus. The spores were about 30×13μm in size usually with two oil droplets. George Grieff kindly identified it as a known but undescribed/unnamed species which he lists as Octospora sp. Bryum-capillare.

Text and photos: Chris Cant

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Lichen

Book reviews, January 2023

Christmas was wet, too wet for doing much outside other than getting damper, so I made a good start on the book pile. Environmental campaigner Guy Shrubsole’s “ The Lost Rainforests of Britain” had a lot of publicity in the autumn: so lots more folk must now be aware of Britain’s rainforests. Job done? Well that must be part of his purpose. The paperback version will tell yet more people about them. And that must be a good thing.

Basically the book is a series of visits to woods, each one an opportunity for Guy to display his growing awareness and to raise issues. I can’t fault his enthusiasm. Indeed, I share it. Once he has discovered rainforests he visits lots; they obviously have a great effect upon him; he becomes a passionate advocate. There are suggested proposals for increasing their area. I can’t really fault them.

I could be picky: what makes something a temperate rainforest?; the species he describes are all too often the same; some of his claims about species and woods and discoveries may be disputed by some; there’s insufficient (to my mind) discussion of the variety of different Atlantic woods. There’s a preponderance (unsurprisingly) of woods near where he lives in south-west England: there’s relatively little about Scottish, Welsh and Cumbrian sites. But, as I said, I’m being picky.

For Cumbrian sites he goes to Johnny’s Wood in Borrowdale and Young Wood near Mungrisdale. I’m not sure those are the best places to go. But that’s from my local lichen perspective and it’s hard to disparage a book where Sticta, Lobaria and others get regular mentions. And maybe we don’t want to encourage too many folk to go near the very special places…

Did it enthuse me? No, but I’m grabbed already. Did it make me want to go to new places? Yes, it’s 30 years since I went to a wood in south west England, so it must be time for a visit soon. Do I recommend it? Well it depends on who you are. If you know a lot about lichens/ bryophytes/ wet woods it may disappoint. And you may be picky. But if you want to broaden the interest, then it may be a good one to suggest for people.

And so to Eoghan Daltun’s “An Irish Atlantic rainforest”. There’s a story here: rebuilding an old house in Dublin; learning about sculpture in Italy; buying a fascinating property in County Cork; fencing it to stop overgrazing killing the woodland. There’s little detail about the wildlife, but great local landscape and social history. The power of a good rainforest to enthuse and interest is revealed. But there’s a series of (to me) blander, less interesting chapters on rewilding and the impoverished state of Ireland’s ecosystems. I knew about that anyway, and maybe I’m spoilt by Tim Robinson’s detailed stories. So I ended up a little disappointed. Not by what Eoghan is doing, which is great, or by what he wants to happen, but by the book. Ho hum.

And then it was Vincent Zonca’s “Lichens: Towards a minimal resistance”, recently translated from the French. It’s a wide ranging tour of art, thought, poetry, prose, biology, ecology, symbiosis, mutualism, philosophy and just about everything else that lichens touch on, or that touch on lichens. The index includes, among those I’ve heard of, John Cage, William Wordsworth, Peter Kropotkin, Salvador Dali, David Hawksworth, Barry Lopez and Rosa Parks. And then there’s all the others.

At times the book is almost unreadable, at times inspiring, at times revealing, at times just pointing at rabbit holes (have a look at: https://www.oscarfurbacken.se/works/urbanlichen).  Rather appropriately, there’s a lot going on, and it’s time consuming to deal with it, even superficially. Have I thought a bit more about things and linkages? Yes. Do I understand it all? No. Does it make me want to know more? Yes. Will I come back to it? Surely, on many occasions. If only to find suitable quotes and inspiration:  “Thinking like lichen allows us to know our ecosystem better, and the everyday environments of our wanderings”(p213). It’s time to go outside in the rain.

Pete Martin

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Bryophyte Lichen Trip report

Glenamara trip report 11-12-22

Bryophyte report below.

Sub-zero licheneering at Glenamara Park, Patterdale

It was well below zero at the carpark with a thin layer of unmelted snow from a previous day on the frozen ground and a thick coating of frost crystals covering everything– surely it wouldn’t be possible to find anything and, more to the point, could we survive the cold? We set off uphill, fat with many layers of clothing, and warmed up slowly. This strategy of a short walk between trees followed by standing around looking at lichens seemed to work and we stayed out until dusk at 4pm.

Glenamara was originally a deer park, dating from the 16th – 17th centuries and covering about 70 hectares of a north-facing upland valley by Ullswater. The National Trust acquired the site in 2002 and shortly afterwards replaced sheep with a small number of hardy cattle as sheep grazing was completely suppressing tree regeneration. Today there are scattered old birch, alder, oak and ash with some hazel, hawthorn and interesting “wild” apple/crab apple trees (Malus sylvestris), but we saw no sign as yet of new trees coming through.

Neil Sanderson surveyed the site for the NT in 2016. He described it as upland pasture woodland and concluded that the only lichen habitat well-represented is that of acid bark communities (Parmelion laevigatae), “due to the past impact of acidifying pollution”, ie acid rain, as well as natural high rainfall acidification (higher than at other sites around Ullswater). He identified only relict base-rich bark (Lobarion pulmonariae) and smooth-bark (Graphidetum scriptae) communities with few species present, as well as small numbers of dry lignum specialists (Calicietum abietinae) – dead wood is well represented at the site.

We had a few of Neil’s locations for interesting crustose species but failed to find them – or perhaps we just didn’t recognise them – so we reverted to the tried and tested method of checking out good-looking trees. Old birch had lovely common acidic bark species, like Ochrolechia tartarea and Mycoblastus sanguinarius, as well as Parmeliopsis hyperopta, a tiny grey foliose species with globose clumps of soredia which isn’t often seen in Cumbria (or most of England). Scattered ash trees had Hypotrachyna taylorensis, new to the Ullswater area, as well as Normandina acroglypta and Catillaria nigroclavata. Diminutive Rinodina sophodes was seen on ash twigs and the equally small apothecia of Dimerella pineti on Malus sylvestris.

A mature oak near the boundary wall at the north edge of the site had a good population of pale Ochrolechia subviridis on the trunk, the granular isidia/soredia showing C+red but no apothecia to be seen. Allan Pentecost had showed us this species in 2019 on an oak lower down beside the playing field, fertile with lovely frosted pruinose discs with isidiate margins. Today’s tree also had Bryobilimbia sanguineoatra, with red-brown apothecia becoming convex with age, growing over moss, as well as a streak of buff-coloured Pyrrhospora quernea.

In the end we had a list of 50-60 species, depending on how the specimens work out, but we wandered over only about a quarter of the site. A return visit as usual may be needed once there’s a thaw and the days are longer.

Text: Caz Walker. Photos: Caz Walker, Chris Cant, Belinda Lloyd, Pete Martin

Glenamara Bryophyte Report

Five hardy bryologists ventured into Glenamara’s frosty wood pasture on a beautiful crisp Sunday morning. The site lies entirely within tetrad NY31X, but straddles all four monads. Due to the conditions the group spent the entire day within monad NY3815. Surprisingly for its location, only 7 species appear to have been recorded for the tetrad in the past so there was a lot to do…

Snow obscured much of the ground so we gravitated towards Hag Beck which was still flowing, and most of the records came from here. After ticking off some commoner grassland species, we started to investigate the beck itself.

Marchantia polymorpha ssp polymorpha proved to be the most ubiquitous thallose liverwort on vertical soil banks all along the beck, with very little Pellia sp present. Plagiochila spinulosa was also found here, together with Fissidens dubius, Trichostomum tenuirostre and Scapania undulata. Platyhypnidium riparoides and Hygrohypnum luridum were common on boulders in the beck itself, together with Thamnobryum alopecuroides, and small bright green patches of Lejeunea patens. Vertical rock above the water-line had cushions of Amphidium mougeotii. Following the beck upstream we found small base-rich areas with Ctenidium molluscum and Tortella tortuosa. Unfortunately, the flushes here were mostly covered with snow, however Sphagnum rusowii and Leiocolea bantriensis were found suggesting that these areas would merit another look. Lunch was necessarily brief due to the penetrating cold, however it did allow us to time to appreciate the stunning views down Ullswater. Two very confiding robins also came to investigate us and hoover up any crumbs. Further up the beck, water was still flowing in several flushes. Green cushions of Blindia acuta were found here together with Palustriella commutata, Hookeria lucens, Rhizomnium punctatum, Cratoneruon filicinum and Riccardia chamaedryfolia. Abundant dead wood was mostly snow-covered, but Riccardia palmata and Tetraphis pellucida were found. By about 2.30 fog started rolling down over Trough Head, so rather than start recording in a new monad we headed down the valley again to find the lichen group. On the way down, Frullania fragilifolia was found on a large Ash tree, its distinctive ‘pear-drops’ scent apparent even in the cold. Having seen no Bazzania trilobata all day, a small patch was located near the beck below the public footpath. As the light was fading,  a tiny Lophozia with red gemmae was found on a very large oak tree on the northern boundary. This could be L. excisa or possibly L. longidens, and has been sent to a referee for determination.

Altogether just over 90 species were recorded, but this would undoubtedly have been higher without the snow. A further visit is clearly required.

Kerry Milligan