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Bryophyte Trip report

Whitbarrow trip bryophyte report 18 February 2024

It was great to welcome a number of new faces amongst the dozen bryologists who assembled in the ample parking area, ready for the fray. The group’s chosen modus operandi, holding field visits which fall on various days of the week including Sundays, is clearly working, allowing people the option to attend when it suits. It surely also helped that Sunday’s forecast was reasonable, and improving, as the day progressed. In truth, the weather was superb for February, everyone enjoying the sunshine, the extensive views from the top of the scarp, and of course the chance to look at mosses without rain!

Whitbarrow is a limestone scar – essentially, the compressed remains of uncounted billions of sea creatures from 350 million years ago, which died and sank to the bottom of the shallow tropical sea, their skeletons of white calcium carbonate slowly turning into Carboniferous Limestone. The tectonic plate drifted from the tropics, to a higher latitude, was uplifted, and now the Scar stands a “mighty” 210m (at highest point) above sea-level, here in Cumbria. Given that it sit on a coastal plain, its modest height nevertheless gives Whitbarrow superb views, both out across the bay, and inland to the higher, more acid hills of the central lakes.

The limestone creates habitats and bryophyte communities which are quite uncommon in Cumbria (have a look at a geological map – there’s not many places where limestone exposures exist). Bryophytes, just like lichens, are sensitive to underlying pH, so the alkaline conditions created on limestone allow a suite of liverwort and moss specialist “calcicoles” to evolve. (Calci = calcium, –colere = “inhabit”, so “things which inhabit calcium-rich areas”). Although soils and seepage water derived from limestone are indeed strongly alkaline, the situation is complicated by the presence of loess (wind deposited sediments) and the development of humus-rich soils in pockets on the limestone, which over time become leached by the glorious Cumbrian rain, and so can become really rather acid in pH. Pockets of acid soil, in a predominantly alkaline habitat, means that the bryophyte communities are rich and varied, and create mosaics where calcicolous and acidophile species can sit side by side. A case in point? We came across two Riccia species, neither of which are all that commonly recorded in Cumbria. Riccias are a group of small liverworts which are colonists of bare, wet, mud – they pack their life cycle into a short time (whilst the mud is wet) and exist as spores in the long dry spells. Riccia beyrichiana grows on acid soils, whereas Riccia sorocarpa is a calcicole. They were two metres apart.

The main objective for the day was to record on Cumbria Wildlife Trust’s Hervey Reserve which includes Lords Seat, the highest point on the scar. Despite being a National Nature Reserve, Site of Special Scientific Interest and Special Area of Conservation, there are several monads on Whitbarrow with surprisingly few records. We climbed the steep path through the wood trying to resist the urge to stop and look at the abundant mosses and liverworts present as this was outside the target square. Once over the stile and into SD4486, the pace slowed as the mosaic of limestone grassland, woodland, and broken rock and scree of the Park Limestone were closely examined. The calcicoles Ctenidium molluscum, Neckera crispa and Tortella tortuosa were conspicuous at almost every stop, and there were fine patches of Flexitrichum (formerly Ditrichum) gracile in places. Closer inspection of some of the Tortella revealed the presence of the uncommon Tortella densa. Acidophiles such as Hylocomium splendens, Hypnum jutlandicum and Polytrichum formosum, and more generalist species such as Thuidium tamariscinum, Pseudoscleropodium purum and Dicranum scoparium were also frequent in the grassland. Close inspection of the ant-hills revealed the identities of numerous small acrocarps (unbranched mosses) including Bryum dichotomum with bulbils visible on the shoot tips, Weissia controversa with abundant capsules, and Barbula convoluta.

After lunch just below Lord’s Seat, we dropped down the dip slope and onto the Urswick Limestone with its distinctive pavements. Here Clare found Rhytidium rugosum growing with (and looking very like) Hypnum cupressiforme var lacunosum (formerly Hypnum lacunosum), and soon we were spotting it everywhere! Rob found Marchantia (formerly Preissia) quadrata growing on the low cliffs, and we also found Tortella nitida here, helpfully growing with T densa and T tortuosa for comparison.  Argles Tarn was an obvious place to visit as a ‘bonus habitat’. The relatively uncommon Campyliadelphus elodes had previously been recorded here and we may have found a small amount growing on pavement near the tarn margin, but this is still to be confirmed. Calliergonella cuspidata and Scorpidium cossonii were also found here. The developing Hazel woodland around the tarn proved to be of interest too, with numerous “corticolous” (on wood) species including the liverworts Myriocoleopsis (formerly Cololejeunia) minutissima, Radula complanata, Metzgeria violacea and Ricardia palmata. On our way back, Campylopus fragilis and Encalypta streptocarpa were added to the list, which at the time of writing stands at 77 species for SD4486.

Report by Gary Lawrence and Kerry Milligan

Photos by Paul Ross (PR), Clare Shaw (CS), Peter Bullard (PB) Gary Lawrence (GL) and Kerry Milligan (KM)

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Lichen Trip report

Whitbarrow trip lichen report 18 February 2024

The last few trips have been dogged by winter deluges. But, mirabile dictu, the forecast for Sunday kept on improving. The rain stopped, the mists cleared and hordes gathered at the Witherslack Hall kennels. Would there be enough parking room? Yes! How many people were there? 23 I think, though only 11 in the lichen party!

As ever, we didn’t get very far very fast. The hawthorns in the first field provided a convenient opportunity to talk form (leafy, crusty etc)  and reproductive features (apothecia, soredia etc). The sun came out; layers were shed. Physcia tenella looked very frilly, and we found a convenient limestone outcrop. Free-living Nostoc was there to be handled, adjacent to the jelly lichen Lathagrium cristatum var. marginale. The latter was in more exposed positions. There was Placynthium nigrum’s blue margin to be admired and the difference between Protoblastenia rupestris and Protoblastenia incrustans could be demonstrated.  A nearby ash had the non-lichenised fungus Hysterium angustatum on it. An hour and a half after starting and we had maybe made it 250 metres.

Bu things speeded up. A bit. The fence by the sports pitch had some Cladonias to admire. The adjacent trees boasted Usnea subfloridana, Hypogymnia tubulosa, and fertile Hypotrachyna revoluta sensu strictu. And then we were into the woods.

A mossy wall with Peltigera membranacea , P. praetextata and Scytinium lichenoides detained us a while. Hazels had Graphis scripta and plentiful Coenogonium cinnabarinum extending in a pink sheen several feet up the stems. Chris found Pachyphiale carneola and there was some Thelotrema lepadinum too. Eventually, in a surge of enthusiasm, we made it up onto the open ground for lunch with a view of Morecambe Bay and Heysham nuclear plant. Was the sea silver, gold, or green? Opinions were divided.

The shattered limestone was possibly less rewarding than hoped, though we managed to pick up some of the common limestone species: Gyalecta jenesis, Romjularia lurida, Collema fuscovirens, Cladonia pocillum, Placidium squamulosum. Sandstone erratics gave lovely mosaics of species we mostly couldn’t identify, though there was a lot of Porpidia tuberculosa. The sad ruins of ash and birch trees gave us a dramatic patch of Trapeliopsis pseudogranulosa.

The afternoon was drawing on and a decision was made to move more quickly. Almost immediately we were detained by hi-vis roundels of Caloplaca flavescens. Sheltered juniper seemed to have more lichens:  one patch had an impressive Parmelioid community. A fenced off area (presumably to stop cattle falling down a hole) had the best Cladonia rangiformis we had seen all afternoon. Deer bounded off across the ripped out pavements.

Soon we were heading down again: there was more Thelotrema lepadinum, and as the longer-lasting members of the party came through the woods, under the cawing ravens and mewling buzzards, we found primroses in bloom. And then, finally, there was an intriguing white sorediate crust on an ash. Chemicals were applied: C+ orangey yellow, K+ yellow, UV+ orange. Is that Lecanora alboflavida? Or Ochrolechia turneri?

“What a good day” someone wrote afterwards. “ It was really fun” someone else said. I enjoyed it immensely. It is good to know others did too.

Text: Pete Martin. Photos: Chris Cant, Pete Martin

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Bryophyte Trip report

Swindale bryophyte trip report 10 December 2023

The weather forecast for 10 December was for non-stop double rain all day, so it was a small and determined group that met at Swindale, three of us looking at bryophytes (Ian, Kerry and Clare) and Caz and Chris looking for lichens. We headed up the valley along the road, wading at times through deep puddles, to reach the bridge by the stepping stones over Swindale Beck. then followed the path to Gouthercrag Gill.

The first stop was by the bridge over the gill. There were many Racomitrium species on rocks near the water, including R. aciculare by the water’s edge, R. lanuginosum, R. heterostichum and R. fasciculare (both checked microscopically); also Polytrichum piliferum and a small cushion of Grimmia donniana with a single capsule. Some good mossy boulders had Barbilophozia attenuata and a small patch of Lophozia sudetica, tentatively identified by the red back to the stem and bilobed leaves. Later, under the dissecting microscope, some nice red perianths were found. A few yards from the beck, a rock face with water running down it had a good range of tufa moss: Amphidium mougeotii, Gymnostomum aeruginosum and Anoectangium aestivum, as well as Palustriella commutata, Conocephalum salebrosum, Preissia quadrata and Fissidens taxifolius. There were healthy patches of Loeskeobryum brevirostre by the beck, and a small amount of Fontinalis antipyretica at the edge of the water.

We headed away from the beck, past flushes where Dichodontium palustre was growing with Philonotis fontana and small amounts of Sphagnum auriculatum and S. inundatum. Scrambling up a slope that proved to be a stable boulder field, we found large amounts of Barbilophozia floerkei. In this promising habitat we hoped to find more Atlantic species such as Anastrepta orcadensis, but the Barbilophozia was predominant. Some of the boulders had large quantities of Gymnomitrion obtusum and some Andreaea rupestris, whilst low crags had cushions of Bartramia pommiformis. At the top of the boulder field, Gouther Crag provided some shelter so we stopped for lunch and a welcome hot drink under a hawthorn which occasionally showered us with drips. Although the crags themselves were dry and largely unpromising, we found some different species on boulders below, including Plagiomnium affine, Plagiothecium denticulatum var. denticulatum, Ptilidium ciliare and a small, narrow-leaved Polytrichum that turned out to be Polytrichastum alpinum. The weather had been showery rather than the unremitting heavy rain that the forecast suggested, but it was still starting to soak into the ends of sleeves and it was a bit on the chilly side to be sitting, so we decided to descend the slope and walk briskly up the valley to look at some hazel woodland. On the way down we passed a large cushion of Tortella tortuosa and an enormous colony of Gymnomitrion obtusum. The hazel woodland produced a few more species including Ulota bruchii, Rhytidiadelphus triquetrus and Frullania tamarisci. It was encouraging to see that the hazels all had abundant young stems as well as older wood. Unfortunately by this stage the rain seemed to be getting heavier and we were getting colder, so we decided to head back. It was a lovely site, quiet and unspoilt, and would definitely be worth revisiting in better weather, perhaps exploring further up the valley and some of the other crags. There was enough variety here to suggest it might warrant further exploration.

Text and photos: Clare Shaw and Kerry Milligan

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Lichen Trip report

Swindale lichen trip report 10 December 2023

Swindale, east of Haweswater

All the lichens looked beautiful, showing their fully hydrated colours. Admittedly, some were actually under water.

Two of us turned up to look at lichens, which is no surprise as this is our local patch so easy to escape home. It’s understandable that for everyone else it wasn’t worth a long drive in order to stand in the rain for several hours.

We’d been here a fair few times and mainly examined the birch, ash and hazel trees beside the path which runs along the bottom of the NW-facing slope below Gouther Crag, so this time we wandered uphill to look at the many boulders which had come down long ago from the crags above. On the way, there was the inevitable lichen-rich drystone wall where, in a dry recess, we saw an intriguing species later identified as Psoronactis dilleniana, new to us. It took a while back at home to work out what this was but luckily it was distinctive (unlike many saxicolous crusts) and Chris nailed it by flicking through the usually poor pictures in Dobson’s guide. The P+ yellow reaction and spore size matched. Interestingly our specimen had a pale but definite K reaction, confirmed by a blue colour under UV, whereas the literature has it as K-.

There was lots of Lecidea lithophila on boulders nearby with apothecia looking very plump and showing their dark brown colour, more visible when wet. Also on the wall was Diploschistes scruposus, looking very yellow, plus other crustose species such as Varicellaria lactea and Lepra aspergilla, separated by easy chemical tests. Unfortunately, wet lichens cause a drop of bleach instantly to spread out which dilutes the reaction (it’s harder or impossible to see) and potentially kills more of the lichen. There are good reasons why licheneering in the rain is a bad idea.

Moving up the vegetated boulder slope, we found Lecanora subcarnea, in a dry-ish NE-facing niche on an outcrop. We’d seen this before nearby in the valley and it was also spotted on the group trip to Burnbanks in May on the sheltered side of a huge boulder. Baeomyces rufus was another eye-catching crust on boulders.

We sheltered for lunch under a dripping holly at the base of a crag, where we found Schismatomma umbrinum and a lovely mosaic of Gyrographa gyrocarpa and Enterographa zonata (both used to be Opegrapha). The party trick of shining a UV torch on a well-covered rock produced spectacular results, Psilolechia lucida glowing orange and chlorophyll in random algae a deep maroon. Also here was Cystocoleus ebeneus, very finely filamentous but looking like black furry patches from a distance.

In order to warm up we wandered on, moving along then down the slope taking in trees on the way. An ash had Mycoblastus sanguinarius, usually found on acidic substrates. A final yellow-orange patch in a sheltered recess on a birch turned out to be Chaenotheca ferruginea with lots of tiny dark pins seen when we examined it closely. The small ornamented spherical spores were nice to see.

This is a great site that needs further visits to do it justice.

Text: Caz Walker. Photos: Chris Cant and Caz Walker

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Lichen Trip report

Blawith Common lichen trip report 14 November 2023

This trip came close to being cancelled when Storm Debi deposited more than 100mm of rain over the 2 preceding days together with gale force winds, but licheneers (and bryologists) are hardy breeds, and we went ahead, given a forecast offering nothing worse than heavy showers. The roads were passable with care, and we had good turnouts for both the lichen and bryophyte wings.

We started with a gentle stroll up the road from Brown Howe car park, admiring the extremely well hydrated state of everything, but particularly the lichens. The first stop was at a roadside rock, liberally crusted with lichens, including a nice patch of Baeomyces rufus.

Baeomyces rufus
Baeomyces rufus

And plenty of Cladonias of which portentosa is usually identifiable…

Cladonia portentosa
Cladonia portentosa

Unlike some others…

Mystery Cladonia
Mystery Cladonia

We didn’t have to go much further to see the star of the show, a veteran Ash with a good population of Pannaria conoplea, a scarce lichen in Cumbria and one of the Lobarion group of Atlantic rainforest species.

Pannaria conoplea ash tree
Pannaria conoplea ash tree
Pannaria conoplea
Pannaria conoplea

It was a pleasant surprise to find it on several nearby willow trees as well since the ash is unlikely to survive much longer even if it escapes the dreaded dieback.

Pannaria conoplea
Pannaria conoplea on a willow

All the common tree lichens were well represented including a nice Usnea cornuta.

Saxicolous lichens perhaps less so due to most of the rocks being well covered with other vegetation.

Usnea cornuta
Usnea cornuta

But this Cladonia was just showing off!

Pete tried to recruit Tip as a lichen sniffer dog but he wasn’t impressed 🙂

Text: John Adams. Photos: John Adams, Pete Martin, Chris Cant.

Chris adds:

In the afternoon, we continued slowly further into woods where there were some more exposed rocky outcrops. Lichens found included Stereocaulon vesuvianum, Psilolechia lucida, Phaeographis smithii, Candelaria concolor and Fuscidea lightfootii.

Phaeographis smithii
Phaeographis smithii
Phaeographis smithii section
Phaeographis smithii section showing broken black exciple underneath hymenium
Candelaria concolor
Candelaria concolor
Fuscidea lightfootii
Fuscidea lightfootii

As the light faded, Peter found an interesting lichen on rock with small black lecideine fruit. Back at base, we thought that it could be Porpidia flavocruenta. Although some characteristics were right, others were not, such as apothecia and hymenium too small and the exciple did not turn red in K. A tricky genus.

Possible Porpidea
Section showing I+ blue reaction spreading down

It was great to find the Pannaria conoplea and see that it was spreading from ash onto willows nearby. However there no other species from the Lobarion community present. That said, there was very little Xanthoria parietina, which was a good sign.

Next time, hopefully we can get further out onto the common.

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Bryophyte Trip report

Blawith Common bryophyte trip report 14 November 2023

On the day before the trip date, the weather was appalling, with strong winds and relentless heavy rain, and the weather forecast was not amazing, so we set off for the day with some trepidation to see how the weather would evolve. There was a lot of standing water on the roads, with quite long stretches of the Burton road submerged, but two of us arrived safely at the meeting point on the A5084, near the small road to Stable Harvey (SD 28934 91049). There was a surprisingly good turnout, with eight people in the bryophyte group.

With the benefit of local knowledge from Rob, we decided to have a quick look at the quarry where some of us were parked. It was indeed a good site and the bare quarry floor was almost entirely carpeted in bryophytes, including Philonotis fontana, small amounts of Dichodontium palustre, Didymodon insulanis, Campylium stellatum, Cratoneuron filicinum, Calliergonella cuspidata and quite a surprising amount of Palustriella commutata (not as regularly pinnate as you would normally expect, but showing all the other features – plentiful rhizoids and paraphylla, curved leaves etc.). We had two beginner bryologists joining the group for the first time, so this was a good opportunity for teaching / revising some common species. The walls of the quarry also had quite a good variety of species: large quantities of Kindbergia praelonga in one corner, also Schistidium crassipilum and some small cushions of fruiting Ptychomitrium polyphyllum. At the far entrance there was further evidence of base-rich substrate, with Tortella tortuosa and Ctenidium molluscum. It was nice to see a good cushion of the tufa moss Gymnostomum aeruginosum, as well as Ptilidium ciliare and a small patch of Scapania compacta.

After a good explore of this area we thought it was time to head for the mires on Blawith common. We splashed up the Stable Harvey road, stopping to admire some lovely patches of the lichen Pannaria conoplea on an old ash (see lichen report!). The lower stretches of the road pass through open woodland with birch and some old ash and oak. Spotting a waterfall just off the road, we headed off to explore. The waterfall had Chionoloma tenuirostre, Metzgeria conjugata and a few nice cushions of Amphidium mougeotii (another tufa moss), but sadly no Jubula hutchinsiae, which has however been recorded in this tetrad. A veteran oak overhanging a rock face had a big cushion of prolifically fruiting Leucobryum at its foot – the capsules (with a white calyptra and distinctive but small bump at the base) pointed to L. glaucum as L. juniperoideum is more rarely seen fruiting and the capsules are slightly different. A sample was examined in case it was L. albidum, but it was confirmed by Tom Blockeel as L. glaucum. Hanging off the rock face, on a mat of decaying vegetative matter, was a small amount of Lophozia incisa. There was also some nice, gemmiferous Lophozia ventricosa on a tree root, Gymocolea inflata on a rock and a potential Rhytidiadelphus subpinnatus. We had lunch near the waterfall, then headed back to the road under a grey sky and light drizzle. Some flat rocks near the road had Polytrichum pilliferum, Racomitrium lanuginosum and Pogonatum urnigerum with Bryum alpinum and Breutelia chrysocoma in seepages. By the road edge there was some gold-tinted Sphagnum inundatum.

A little way up the road, we finally reached the mire, which at first was rather poor, mainly Molinia and Myrica gale with only small amounts of Sphagnum beneath. We collected a couple of Sphagnum species, then headed across the road to a better site, where there were cushions of Polytrichum strictum and Aulocomium palustre, as well as several Sphagnums: S. capillifolium subsp. capillifolium, S. auriculatum, S. fallax, S. inundatum, S. palustre, S. papillosum and S. russowii. There were patches of Odontoschisma Sphagni and one patch of Mylia anomala, with only a few shoots showing clusters of gemmae after the heavy rain. Straminergon stramineum was poking its head out of some of the mounds. A small escarpment of rocks above the mire had Andreaea rothii and Ptilidium ciliare.

A cold wind was starting to blow, so we decided to beat a retreat. It was an enjoyable day, and undoubtedly far better than any of us had been expecting.

Text and photos: Clare Shaw

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Bryophyte Trip report

Duddon Valley bryophyte trip report 15 October 2023

For a change the sun was shining and the winds light for our October meeting at High Wallowbarrow Farm near Seathwaite in the Duddon Valley. The farm is unusual in having large areas of ancient semi-natural woodland, most of which lies within the Duddon Valley Woodlands Site of Special Scientific Interest. Whilst bryophytes have been quite well recorded in parts of the SSSI, other areas appear to have been little visited; monad SD2196 immediately west of the farmhouse  has records for only three species, whilst the adjacent monad (SD2296) has a list of 189 species. We decided to focus the morning’s efforts in SD2196 and visit the spectacular Wallowbarrow Gorge in SD2296 in the afternoon.

After a brief introduction, we made our way to Wallowbarrow Coppice, an area of upland oakwood on a steep west facing slope with frequent birch and hazel. An abundance of common mosses such as Thuidium tamariscinum, Isothecium myosuroides, Rhytidiadelphus loreus and Polytrichum formosum covered the woodland floor, rocks and tree bases, with the liverworts Scapania nemorea, Scapania gracilis, Diplophyllum albicans also abundant. More unusual species included Rhabdoweisia crenulata, Orthocaulis (Barbilophozia) atlantica, Tritomaria exsectiformis, and Scapania umbrosa. A quick foray into a small mire above the wood added several Sphagna including Sphagnum papillosum and S rubellum, together with Polytricum strictum and Aulocomium palustre. An unusual Racomitrium on rocks nearby was later confirmed as R obtusum. This was formerly considered a variety or subspecies of R. heterostichum and there are as yet very few records of it in Cumbria, although it is likely to be quite widespread. Whilst we did not re-find Syzigiella (Jamesoniella) autumnalis, Grimmia ramondii or Ptychomitrium polyphyllum which had previously been found in SD2196, 62 new species were recorded. We could probably have spent the whole day in this monad and identified a lot more, but after lunch we headed into SD2296 to visit Crag End Wood and Wallowbarrow Gorge. Quickly ticking off many of the common bryophytes already encountered in the morning, we soon found some new species including Diphyscum foliosum and Hyocomium armoricum on the river banks, Sciuro-hypnum plumosum and Marsupella aquatica on rocks in the river, Amphidium mougeotii and Saccogyna viticulosa on crags, and Dicranodontium denudatum on dead wood. Towards the end of the day, the rain came in and we retreated. In total 83 bryophytes were recorded across the two monads. Thanks to Chris at High Wallowbarrow Farm for access and to everyone who participated in what was a very enjoyable day.

Text: Kerry Milligan. Photos: Kerry Milligan, Clare Shaw, Paul Ross.

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Lichen Trip report

Duddon Valley lichen trip report 15 October 2023 

A large group met at High Wallowbarrow Farm in the Duddon Valley on a sunny autumn day. There are few lichen records here, monad SD2296 having 34 mainly from 1970 (Brian Coppins and Francis Rose). We started by looking in the next square to the west which was unrecorded. This is a formerly coppiced wooded slope where alder, birch, hazel and oak grow amongst huge mossy boulders and outcrops.

It was slow going as every tree and rock needed examination, with lots of lichen chat. Many participants were complete beginners while others had come on trips earlier in the year and remembered enough to have a go at identification – and could help those who knew less. Normandina pulchella on several hazel trees proved popular, as well as discussions around various Cladonia species. A large boulder had a granular yellow-green and orange patch of Trapeliopsis pseudogranulosa as well as some crustose Pertusaria species to which chemicals were applied in order to confirm the identification with a colour change.

John Douglass, a lichenologist based in Scotland, was with us and helped point things out to everyone, including Micarea alabastrites with flat, white apothecia on a mossy alder and crustose species, such as Rhizocarpon infernulum f. sylvaticum, Porina lectissima on a mossy boulder high in the wood and both Trapelia glebulosa and T involuta growing side by side on a massive boulder beside the path near the farm. The latter is a distinct species but has been confused with T glebulosa in the past. Also here were Arthrorhaphis citrinella, easily identifiable with a bright yellow-green granular thallus and, in this case, fertile with black apothecia, as well as Placopsis lambii, a crustose lichen which has the appearance of lobes around the edge as well as flat patches of grey soredia, making it very recognisable. Nearby a birch had tiny neat pale yellowish rosettes of Parmeliopsis ambigua, with soredia arranged in globose soralia on the narrow lobes. Furrowed lirellae of Graphis elegans were on the same tree.

After lunch John headed to the river to search for aquatic lichens (his specialism) on damp rocks beside the river, finding Porina rivalis, new to VC70, as well as commoner species like Massalongia carnosa, Ephebe lanata and Ionaspis lacustris.  The rest of us moved slowly along a dry stone wall, finding lots of lichen interest. Geoffrey enjoyed exploring the species there with Carole and Paul, taking the time to look closely at some tiny features. These walls can be very good for lichens and this was no exception – many little habitat niches supported plenty of common species as well as Stereocaulon pileatum and some good Cladonia species, such as the red-fruited C diversa and C bellidiflora. Mature oak and ash in the pasture had a range of foliose lichens and a hazel at the woodland entrance had the typical smooth bark species Pertusaria leioplaca and Arthonia radiata.

A final mossy boulder had good Cladonia which deserved close inspection, Cladonia caespiticia being the most obvious as it was fertile with tiny pink mushroom-like structures and minute dark pycnidia on the squamules. Also here was Cladonia squamosa showing pink brown apothecia – it’s not often seen fertile.

Overall there was something for everyone here but the feeling was that quite a lot was missing – there wasn’t the range of lichens or bryophytes we might expect at a good site. Kerry pointed out that the area had been intensively exploited in the past – heavily grazed, coppiced, trees felled for firewood etc – not to mention over 100 years of pollution from the industrial SW Cumbrian coast, all of which explains what we see, or don’t see, today. The recovery of the biodiversity may take many years.

Text: Caz Walker: Photos: Caz Walker, Chris Cant, Geoffrey Haigh, John Douglass

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Bryophyte Trip report

Mallerstang Bryophyte report 14 September 2023

As with any bryology outing, it’s easy to get distracted by the surroundings of the car park. Some awkward road closures and traffic spread out arrivals, so a brief look around Pendragon castle yielded a light green Zygodon growing on an old ash – Zygodon rupestris perhaps? Alas, the longitudinal cell walls on its gemmae confirmed it as the more-common Zygodon viridissimus.

Once all three bryologists were present, we set off for the Mallerstang crags aiming to focus our efforts on grid square NY7902 which curiously lacked any bryophyte records. After confirming the presence of Rhytidiadelphus squarrosus ambitiously (presumptively?) identified by Judith from 800m away in the car park, we found a typical assembly of upland acid grassland mosses and the first few of what would be a good day for SphagnumSphagnum capillifolium, S. subnitens and S. russowii.

Our first liverwort of the day was Tritomaria quinquedentata growing among wefts of Pleurozium and Pseudoscleropodium, quickly followed by Ptilidium ciliare, Orthocaulis attenuatus and a larger gemmiferous Orthocaulis (confirmed later as O. atlanticus) on the grassy margins of exposed rocks. The shaded bases of the larger rocks were occupied by Diplophyllum albicans, Scapania gracilis and the occasional large stand of a Cephalozia, later confirmed by Kerry to be C. bicuspidata.

As we advanced further towards the crags, we crossed several rush-dominated flushes, adding six more sphagna to our list – S. squarrosum, S. auriculatum, S. palustre, S. fallax, S. angustifolium and S. teres. We also added Polytrichum commune whose family was well represented on the day, including P. piliferum, P. strictum, P. formosum, Polytrichastrum alpinum, Pogonatum urnigerum and P. aloides.

We passed a few more damp patches adding Sphagnum papillosum and S. rubellum to our list, and our arrival at the crags was further delayed by a stream with typical riparian bryophytes – Brachythecium plumosum, Racomitrium aciculare, R. aquaticum, Scapania undulata and a Pellia species. The flushed edges boasted Philonotis fontana and Dichodontium pellucidum with scattered patches of Riccardia chamaedryfolia.

It was after 2pm when we reached the crags where we pondered a slender Campylium, later identified as C. portensum with leaves <2mm long, large alar patches and abruptly tapering leaves. The flushed crags were adorned with flows of the usual suspects – Philonotis fontana, Amphidium mougeotii, Palustriella commutata and Breutelia chrysocoma. The diversity expanded with Pohlia wahlenbergii, Bryum julaceum, Aneura pinguis, Pellia endiviifolia with ample branchlets, Blindia acuta, Jungermannia atrovirens, Hookeria lucens and Gymnostomum aeruginosum.

Our journey back to the carpark included a brief stop around some boulders with Calypogeia arguta, Schistidium crassipilum and Orthotrichum diaphanum joining our list. Further down, a quick look around a streamside limestone exposure yielded little more of interest but hiding in turf overhang was Sphagnum girgensohnii, our twelfth and final Sphagnum of the day.

Text and photos: Paul Ross

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Lichen Trip report

Mallerstang Lichen Report 14 September 2023

Lindrigg Scars, Mallerstang

A day of fine weather following overnight rain gave perfect conditions as the lichens were well hydrated and colourful but not too sodden. We parked opposite 12th century Pendragon Castle and had a quick look at the walls and adjacent ash trees, sadly badly affected by die-back, then plodded up hill towards the gritstone cliffs at about 600m on the eastern skyline. Once above the fell wall, isolated rocks started to appear in the grassy slopes with proper boulder fields higher up. Some of these rocks had swards of lichens amongst moss, ranging from larger leafy species like Parmelia saxatilis, Hypogymnia physodes and Platismatia glauca to smaller crustose species including plentiful Rhizocarpon, Porpidia, Pertusaria and Fuscidea, as well as colourful Ophioparma ventosa. One patch of Parmelia saxatilis was being consumed by the lichenicolous fungus Marchandiomyces corallinus resulting in pale necrotic areas scattered with pink bulbils. Nearby brown P omphalodes had a different LF with black fruiting bodies bursting through the cortex – there’s a specimen to examine. There wasn’t a wide range of Cladonia species, probably because there was little exposed peat with generally dense grasses covering the ground. A niche for these can be found, however, along the turf-rock boundary where there is less competition and especially on rock surfaces where there is a “biological crust” of dead bryophytes, lichens and algae. Here we found red-fruited Cladonia diversa and C polydactyla, the brown pointed podetia of Cladonia furcata, the elegant Cladonia gracilis and glossy dark chestnut Cetraria aculeata with tiny white oval pseudocyphellae immersed in the branches. This is a fruticose (bushy) lichen, as is Bryoria fuscescens, found amongst moss and other lichens on the north side of a large boulder. One Cladonia new to us was C luteoalba, the undersides of the squamules densely tomentose/fluffy. This species is supposed to start life on other Cladonia species, presumably pinching their algae.

Some of the party expanded their horizons to include fungus as this is the start of the waxcap season. Judith found the large pink Ballerina waxcap, Porpolomopsis calyptriformis, and a few others were seen later.

There were fewer species on the vertical gritstone of Lindrigg Scars but, associated with thin beds of siltstone, were a couple of small aspen trees giving us the only records of corticolous species – Lecidella elaeochroma and Arthonia radiata. The aspen may have got there as a seed (though it’s not often fertile in the UK and there were no other nearby trees) and has survived out of reach of browsers – unless it’s a surviving sucker from an original tree now gone. On the rock face here were patches of brown Parmelia discordans, similar to the commoner P omphalodes but without obvious pseudocyphellae and with a K- medulla (P omphalodes is K+ orange). Nearby, we found Lichenomphalia alpina where peat was hanging over an outcrop. There was some doubt as the mushroom fruiting bodies were a paler yellow than we’d seen before, but the caps were smaller in diameter than the commoner L umbellifera, which has a browner cap with a wavy margin and more striate. Both share the same thallus of shiny dark green globules of clustered algae. Lichenomphalia velutina was also mentioned as a possibility but that also has a brownish cap. A short distance further on there was another patch of L alpina, this time with tiny developing buttons which were a deeper yellow.

After exploring southward along the edge we headed down, finding a small area of lovely heath on the steep slope below with low-growing bilberry, heather and moss. Here Chris found one small clump of Cetraria islandica as well as the only Umbilicaria species we’d seen, the common U polyphylla. The sward contained plentiful “reindeer” lichens – Cladonia arbuscula, C ciliata and C uncialis.

The monad NY7902, covering the main part of Lindrigg Scars, has 95 existing records. Most of the lichens we saw are covered by this list but with a few new ones so taking the total count for the square over 100. We went into the next monad to the south which seemed to have no records and were able to add over 50 lichens to that square. Overall none of our records had rarity value – the tricky crustose species Porpidia melinodes and P flavocruenta (if confirmed when we have time to look at the specimen) were the only ones less common – but, as Brian Coppins says, it’s important to record the common background species to get a full picture of the habitat. We’re certainly good at that!

Text: Caz Walker. Photos: Chris Cant and Caz Walker